About the Authors

Jonathan L. Zelner

jzelner@princeton.edu

Affiliations Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Benjamin A. Lopman

Affiliation Epidemiology Branch, Division of Viral Diseases, National Center for Immunization and Respiratory Diseases, Centers for Disease Control and Prevention, Atlanta, Georgia, United States of America

Aron J. Hall

Affiliation Epidemiology Branch, Division of Viral Diseases, National Center for Immunization and Respiratory Diseases, Centers for Disease Control and Prevention, Atlanta, Georgia, United States of America

Sebastien Ballesteros

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Bryan T. Grenfell

Affiliations Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JZ BL AH. Performed the experiments: JZ. Analyzed the data: JZ. Contributed reagents/materials/analysis tools: SB. Wrote the paper: JZ BL AH BG.