About the Authors

Theresa M. Nogeire

tnogeire@gmail.com

Affiliation School of Environmental and Forest Sciences, University of Washington, Seattle, Washington United States of America

Frank W. Davis

Affiliation Bren School of Environmental Science and Management, University of California Santa Barbara, Santa Barbara, California, United States of America

Jennifer M. Duggan

Affiliation School of Environmental and Forest Sciences, University of Washington, Seattle, Washington United States of America

Kevin R. Crooks

Affiliation Department of Fish, Wildlife, and Conservation Biology, Colorado State University, Fort Collins, Colorado, United States of America

Erin E. Boydston

Affiliation U.S. Geological Survey, Western Ecological Research Center, Thousand Oaks, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FD TN. Performed the experiments: TN. Analyzed the data: TN JD FD KC EB. Contributed reagents/materials/analysis tools: FD TN. Wrote the paper: TN FD EB JD KC.