About the Authors

Collette M. Thogerson

Affiliation Ecological Services, U.S. Fish & Wildlife Service, Arlington, Virginia, United States of America

Colleen M. Brady

Affiliation Department of Animal Sciences, Purdue University, West Lafayette, Indiana, United States of America

Richard D. Howard

Affiliation Department of Biological Sciences, Purdue University, West Lafayette, Indiana, United States of America

Georgia J. Mason

Affiliation Department of Animal and Poultry Science, University of Guelph, Ontario, Canada

Edmond A. Pajor

Affiliation Veterinary Medicine, University of Calgary, Alberta, Canada

Greg A. Vicino

Affiliation Collections Husbandry Science, San Diego Zoo Global, San Diego, California, United States of America

Joseph P. Garner

jgarner@stanford.edu

Affiliation Department of Comparative Medicine, and by Courtesy, Department of Psychiatry and Behavioral Sciences, Stanford University, Stanford, California, United States of America

Competing Interests

The authors have read the journal’s policy declare that GAV has the following conflicts: GAV is employed by SDZG. SDZG had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CMT GAV JPG. Performed the experiments: CMT JPG. Analyzed the data: CMT RDH JPG. Contributed reagents/materials/analysis tools: GAV. Wrote the paper: CMT GJM JPG. Contributed to intellecutal development of the project: EAP RDH CMB.