About the Authors

Fiona C. Ingleby

Affiliations Centre for Ecology and Conservation, School of Biosciences, University of Exeter, Cornwall Campus, Tremough, Penryn, United Kingdom, University of Sussex, John Maynard Smith Building, Falmer, Brighton, United Kingdom

John Hunt

Affiliation Centre for Ecology and Conservation, School of Biosciences, University of Exeter, Cornwall Campus, Tremough, Penryn, United Kingdom

David J. Hosken

D.J.Hosken@exeter.ac.uk

Affiliation Centre for Ecology and Conservation, School of Biosciences, University of Exeter, Cornwall Campus, Tremough, Penryn, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FCI JH DJH. Performed the experiments: FCI. Analyzed the data: FCI. Contributed reagents/materials/analysis tools: JH DJH. Wrote the paper: FCI DJH JH.