About the Authors

Qiang Chen

Contributed equally to this work with: Qiang Chen, Yufang Ma

Affiliation School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China

Yufang Ma

Contributed equally to this work with: Qiang Chen, Yufang Ma

Affiliation School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China

Yumei Yang

Affiliations School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China, Jiangsu Normal University, Xuzhou, China

Zhenliang Chen

Affiliation School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China

Rongrong Liao

Affiliation School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China

Xiaoxian Xie

Affiliation School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China

Zhen Wang

Affiliation School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China

Pengfei He

Affiliation School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China

Yingying Tu

Affiliation Shanhai Genome Biotechnology Company, Shanghai, China

Xiangzhe Zhang

Affiliation School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China

Changsuo Yang

Affiliation National Research Centre on Poultry Engineering Technology, Shanghai Academy of Agricultural Science, Shanghai, China

Hongjie Yang

Affiliation National Animal Husbandry Services, Beijng, China

Fuqing Yu

Affiliation National Animal Husbandry Services, Beijng, China

Youmin Zheng

Affiliation National Animal Husbandry Services, Beijng, China

Zhiwu Zhang

Affiliation Institute for Genomic Diversity, Cornell University, Ithaca, New York, United States of America

Qishan Wang

panyuchun1963@yahoo.com.cn (YP); wangqishan@sjtu.edu.cn (QW)

Affiliation School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China

Yuchun Pan

panyuchun1963@yahoo.com.cn (YP); wangqishan@sjtu.edu.cn (QW)

Affiliations School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, China, Shanghai Key Laboratory of Veterinary Biotechology, Shanghai, China

Competing Interests

One of the authors is employed by a commercial company (Shanhai Genome Biotechnology Company, Shanghai, China). The authors declare that this does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: YP QW. Performed the experiments: QC ZZ ZC RL XX CY HY XZ. Analyzed the data: QC YY YM ZW PH YT. Contributed reagents/materials/analysis tools: HY FY YZ ZZ. Wrote the paper: QC QW.