About the Authors

Lindsey E. Carlin

Affiliations Interdisciplinary Graduate Program in Immunology, University of Iowa, Iowa City, Iowa, United States of America, The Department of Pathology, University of Iowa, Iowa City, Iowa, United States of America

Natalya V. Guseva

Affiliation The Department of Pathology, University of Iowa, Iowa City, Iowa, United States of America

Michael R. Shey

Affiliation The Iowa City VA Medical Center, University of Iowa, Iowa City, Iowa, United States of America

Zuhair K. Ballas

Affiliations Interdisciplinary Graduate Program in Immunology, University of Iowa, Iowa City, Iowa, United States of America, The Department of Internal Medicine, University of Iowa, Iowa City, Iowa, United States of America, The Iowa City VA Medical Center, University of Iowa, Iowa City, Iowa, United States of America

Jonathan W. Heusel

jon-heusel@uiowa.edu

Affiliations Interdisciplinary Graduate Program in Immunology, University of Iowa, Iowa City, Iowa, United States of America, The Department of Pathology, University of Iowa, Iowa City, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JWH. Performed the experiments: LEC NVG MRS. Analyzed the data: LEC MRS. Contributed reagents/materials/analysis tools: ZKB. Wrote the paper: LEC JWH.