About the Authors

Nicholas J. Kaplinsky

nkaplin1@swarthmore.edu

Affiliation Department of Biology, Swarthmore College, Swarthmore, Pennsylvania, United States of America

Scott F. Gilbert

Affiliation Department of Biology, Swarthmore College, Swarthmore, Pennsylvania, United States of America

Judith Cebra-Thomas

Affiliation Department of Biology, Millersville University, Millersville, Pennsylvania, United States of America

Kersti Lilleväli

Affiliations Department of Developmental biology, Institute of Molecular and Cell Biology, University of Tartu, Tartu, Estonia, Department of Physiology, Faculty of Medicine, University of Tartu, Tartu, Estonia

Merly Saare

Affiliation Department of Developmental biology, Institute of Molecular and Cell Biology, University of Tartu, Tartu, Estonia

Eric Y. Chang

Affiliation Department of Biology, Swarthmore College, Swarthmore, Pennsylvania, United States of America

Hannah E. Edelman

Affiliation Department of Biology, Swarthmore College, Swarthmore, Pennsylvania, United States of America

Melissa A. Frick

Affiliation Department of Biology, Swarthmore College, Swarthmore, Pennsylvania, United States of America

Yin Guan

Affiliation Department of Biology, Swarthmore College, Swarthmore, Pennsylvania, United States of America

Rebecca M. Hammond

Affiliation Department of Biology, Swarthmore College, Swarthmore, Pennsylvania, United States of America

Nicholas H. Hampilos

Affiliation Department of Biology, Swarthmore College, Swarthmore, Pennsylvania, United States of America

David S. B. Opoku

Affiliation Department of Biology, Swarthmore College, Swarthmore, Pennsylvania, United States of America

Karim Sariahmed

Affiliation Department of Biology, Swarthmore College, Swarthmore, Pennsylvania, United States of America

Eric A. Sherman

Affiliation Department of Biology, Swarthmore College, Swarthmore, Pennsylvania, United States of America

Ray Watson

Affiliation Department of Biology, Swarthmore College, Swarthmore, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NJK SFG JC KL MS EYC HEE MAF YG RMH NHH DSO KS EAS RW. Performed the experiments: NJK JC KL MS EYC HEE MAF YG RMH NHH DSO KS EAS RW. Analyzed the data: NJK SFG JC KL MS EYC HEE MAF YG RMH NHH DSO KS EAS RW. Contributed reagents/materials/analysis tools: NJK JC KL MS. Wrote the paper: NJK SFG JC KL MS EYC HEE MAF YG RMH NHH DSO KS EAS RW.