About the Authors

Ruei-Min Lu

Contributed equally to this work with: Ruei-Min Lu, Min-Shan Chen, De-Kuan Chang

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Min-Shan Chen

Contributed equally to this work with: Ruei-Min Lu, Min-Shan Chen, De-Kuan Chang

Affiliations Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan, Graduate Institute of Oral Biology, College of Medicine, National Taiwan University, Taipei, Taiwan

De-Kuan Chang

Contributed equally to this work with: Ruei-Min Lu, Min-Shan Chen, De-Kuan Chang

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Chien-Yu Chiu

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Wei-Chuan Lin

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Shin-Long Yan

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Yi-Ping Wang

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Yuan-Sung Kuo

Affiliation Department of Surgery, National Taiwan University Hospital, National Taiwan University, Taipei, Taiwan

Chen-Yun Yeh

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Albert Lo

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Han-Chung Wu

hcw0928@gate.sinica.edu.tw

Affiliations Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan, Graduate Institute of Oral Biology, College of Medicine, National Taiwan University, Taipei, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HCW RML. Performed the experiments: RML MSC DKC CYC WCL CYY YPW AL YSK. Analyzed the data: MSC DKC SLY. Wrote the paper: HCW RML.