About the Authors

Christian Hoffmann

Affiliations Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America, Instituto de Ciências Biológicas, Universidade Federal de Goiás, Goiania, Goiás, Brazil

Serena Dollive

Affiliation Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Stephanie Grunberg

Affiliation Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Jun Chen

Affiliation Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America

Hongzhe Li

Affiliations Center for Clinical Epidemiology and Biostatistics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America, Department of Biostatistics and Epidemiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Gary D. Wu

gdwu@mail.med.upenn.edu (GDW); lewisjd@mail.med.upenn.edu (JDL); bushman@mail.med.upenn.edu (FDB)

Affiliation Division of Gastroenterology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

James D. Lewis

gdwu@mail.med.upenn.edu (GDW); lewisjd@mail.med.upenn.edu (JDL); bushman@mail.med.upenn.edu (FDB)

Affiliations Center for Clinical Epidemiology and Biostatistics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America, Department of Biostatistics and Epidemiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Frederic D. Bushman

gdwu@mail.med.upenn.edu (GDW); lewisjd@mail.med.upenn.edu (JDL); bushman@mail.med.upenn.edu (FDB)

Affiliation Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CH GDW JDL FDB. Performed the experiments: CH SD SG. Analyzed the data: CH JC HL. Wrote the paper: CH FDB.