About the Authors

Alessio Boattini

Contributed equally to this work with: Alessio Boattini, Begoña Martinez-Cruz

Affiliation Laboratorio di Antropologia Molecolare, Dipartimento di Scienze Biologiche, Geologiche e Ambientali, Università di Bologna, Bologna, Italy

Begoña Martinez-Cruz

Contributed equally to this work with: Alessio Boattini, Begoña Martinez-Cruz

Affiliation Institut de Biologia Evolutiva (CSIC-UPF), Departament de Ciències de la Salut i de la Vida, Universitat Pompeu Fabra, Barcelona, Spain

Stefania Sarno

Affiliation Laboratorio di Antropologia Molecolare, Dipartimento di Scienze Biologiche, Geologiche e Ambientali, Università di Bologna, Bologna, Italy

Christine Harmant

Affiliations Institut Pasteur, Human Evolutionary Genetics Unit, Department of Genomes and Genetics, Paris, France, Centre National de la Recherche Scientifique, Paris, France

Antonella Useli

Affiliation Dipartimento di Scienze della Natura e del Territorio, Università di Sassari, Sassari, Italy

Paula Sanz

Affiliation Institut de Biologia Evolutiva (CSIC-UPF), Departament de Ciències de la Salut i de la Vida, Universitat Pompeu Fabra, Barcelona, Spain

Daniele Yang-Yao

Affiliation Laboratorio di Antropologia Molecolare, Dipartimento di Scienze Biologiche, Geologiche e Ambientali, Università di Bologna, Bologna, Italy

Jeremy Manry

Affiliations Institut Pasteur, Human Evolutionary Genetics Unit, Department of Genomes and Genetics, Paris, France, Centre National de la Recherche Scientifique, Paris, France

Graziella Ciani

Affiliation Laboratorio di Antropologia Molecolare, Dipartimento di Scienze Biologiche, Geologiche e Ambientali, Università di Bologna, Bologna, Italy

Donata Luiselli

Affiliation Laboratorio di Antropologia Molecolare, Dipartimento di Scienze Biologiche, Geologiche e Ambientali, Università di Bologna, Bologna, Italy

Lluis Quintana-Murci

Affiliations Institut Pasteur, Human Evolutionary Genetics Unit, Department of Genomes and Genetics, Paris, France, Centre National de la Recherche Scientifique, Paris, France

David Comas

david.comas@upf.edu (DC); davide.pettener@unibo.it (DP)

Affiliation Institut de Biologia Evolutiva (CSIC-UPF), Departament de Ciències de la Salut i de la Vida, Universitat Pompeu Fabra, Barcelona, Spain

Davide Pettener

david.comas@upf.edu (DC); davide.pettener@unibo.it (DP)

Affiliation Laboratorio di Antropologia Molecolare, Dipartimento di Scienze Biologiche, Geologiche e Ambientali, Università di Bologna, Bologna, Italy

the Genographic Consortium

Membership of the Genographic Consortium is provided in the Acknowledgments.

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DP DL DC. Performed the experiments: BMC AU DYY SS GC CH JM LQM PS. Analyzed the data: AB SS BMC AU. Contributed reagents/materials/analysis tools: DL DP DC LQM. Wrote the paper: AB BMC SS DC AU. Performed field work, sampling design and collection: AB DYY AU DL DP.