About the Authors

Chuanfu Zhang

Contributed equally to this work with: Chuanfu Zhang, Shaofu Qiu, Yong Wang, Lihua Qi, Rongzhang Hao, Xuelin Liu, Yun Shi, Xiaofeng Hu, Daizhi An, Zhenjun Li, Peng Li, Ligui Wang

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Shaofu Qiu

Contributed equally to this work with: Chuanfu Zhang, Shaofu Qiu, Yong Wang, Lihua Qi, Rongzhang Hao, Xuelin Liu, Yun Shi, Xiaofeng Hu, Daizhi An, Zhenjun Li, Peng Li, Ligui Wang

hongbinsong@263.net (HBS); qiushf0613@hotmail.com (SFQ)

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Yong Wang

Contributed equally to this work with: Chuanfu Zhang, Shaofu Qiu, Yong Wang, Lihua Qi, Rongzhang Hao, Xuelin Liu, Yun Shi, Xiaofeng Hu, Daizhi An, Zhenjun Li, Peng Li, Ligui Wang

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Lihua Qi

Contributed equally to this work with: Chuanfu Zhang, Shaofu Qiu, Yong Wang, Lihua Qi, Rongzhang Hao, Xuelin Liu, Yun Shi, Xiaofeng Hu, Daizhi An, Zhenjun Li, Peng Li, Ligui Wang

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Rongzhang Hao

Contributed equally to this work with: Chuanfu Zhang, Shaofu Qiu, Yong Wang, Lihua Qi, Rongzhang Hao, Xuelin Liu, Yun Shi, Xiaofeng Hu, Daizhi An, Zhenjun Li, Peng Li, Ligui Wang

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Xuelin Liu

Contributed equally to this work with: Chuanfu Zhang, Shaofu Qiu, Yong Wang, Lihua Qi, Rongzhang Hao, Xuelin Liu, Yun Shi, Xiaofeng Hu, Daizhi An, Zhenjun Li, Peng Li, Ligui Wang

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Yun Shi

Contributed equally to this work with: Chuanfu Zhang, Shaofu Qiu, Yong Wang, Lihua Qi, Rongzhang Hao, Xuelin Liu, Yun Shi, Xiaofeng Hu, Daizhi An, Zhenjun Li, Peng Li, Ligui Wang

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Xiaofeng Hu

Contributed equally to this work with: Chuanfu Zhang, Shaofu Qiu, Yong Wang, Lihua Qi, Rongzhang Hao, Xuelin Liu, Yun Shi, Xiaofeng Hu, Daizhi An, Zhenjun Li, Peng Li, Ligui Wang

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Daizhi An

Contributed equally to this work with: Chuanfu Zhang, Shaofu Qiu, Yong Wang, Lihua Qi, Rongzhang Hao, Xuelin Liu, Yun Shi, Xiaofeng Hu, Daizhi An, Zhenjun Li, Peng Li, Ligui Wang

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Zhenjun Li

Contributed equally to this work with: Chuanfu Zhang, Shaofu Qiu, Yong Wang, Lihua Qi, Rongzhang Hao, Xuelin Liu, Yun Shi, Xiaofeng Hu, Daizhi An, Zhenjun Li, Peng Li, Ligui Wang

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, China Center of Disease Control and Prevention, Beijing, People’s Republic of China

Peng Li

Contributed equally to this work with: Chuanfu Zhang, Shaofu Qiu, Yong Wang, Lihua Qi, Rongzhang Hao, Xuelin Liu, Yun Shi, Xiaofeng Hu, Daizhi An, Zhenjun Li, Peng Li, Ligui Wang

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Ligui Wang

Contributed equally to this work with: Chuanfu Zhang, Shaofu Qiu, Yong Wang, Lihua Qi, Rongzhang Hao, Xuelin Liu, Yun Shi, Xiaofeng Hu, Daizhi An, Zhenjun Li, Peng Li, Ligui Wang

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Jiajun Cui

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Pan Wang

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Liuyu Huang

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

John D. Klena

Affiliation United States Centers for Disease Control and Prevention, China –US Collaborative Program on Emerging and Re-emerging Infectious Diseases, Beijing, People’s Republic of China

Hongbin Song

hongbinsong@263.net (HBS); qiushf0613@hotmail.com (SFQ)

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HBS CFZ SFQ YW RZH. Performed the experiments: CFZ XLL YS XFH DZA LHQ ZJL PW. Analyzed the data: CFZ JJC PL LGW LYH. Contributed reagents/materials/analysis tools: YS XFH DZA LHQ. Wrote the paper: CFZ SFQ HBS JDK.