About the Authors

Janet Monge

Affiliation University of Pennsylvania Museum, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Morrie Kricun

Affiliation Department of Radiology, University of Pennsylvania Museum, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Jakov Radovčić

Affiliation Croatian Natural History Museum, Zagreb, Croatia

Davorka Radovčić

Affiliation Croatian Natural History Museum, Zagreb, Croatia

Alan Mann

Affiliation Department of Anthropology, Princeton University, Princeton, New Jersey, United States of America

David W. Frayer

frayer@ku.edu

Affiliation Department of Anthropology, University of Kansas, Lawrence, Kansas, United States of America

Competing Interests

The authors state that Monge and Frayer are Academic Editors. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JM MK JR AM DWF. Performed the experiments: JM MK DR AM. Analyzed the data: MK DR AM. Contributed reagents/materials/analysis tools: JM MK DR. Wrote the paper: JM MK AM DWF.