About the Authors

Betty R. Liu

Affiliation Department of Natural Resources and Environmental Studies, National Dong Hwa University, Hualien, Taiwan

Ji-Sing Liou

Affiliation Department of Natural Resources and Environmental Studies, National Dong Hwa University, Hualien, Taiwan

Yue-Wern Huang

Affiliation Department of Biological Sciences, Missouri University of Science and Technology, Rolla, Missouri, United States of America

Robert S. Aronstam

Affiliation Department of Biological Sciences, Missouri University of Science and Technology, Rolla, Missouri, United States of America

Han-Jung Lee

hjlee@mail.ndhu.edu.tw

Affiliation Department of Natural Resources and Environmental Studies, National Dong Hwa University, Hualien, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HJL. Performed the experiments: BRL JSL. Analyzed the data: YWH RSA HJL. Wrote the paper: HJL BRL YWH RSA.