About the Authors

Angela M. White

angelawhite@fs.fed.us

Affiliation Conservation of Biodiversity Group, Pacific Southwest Research Station, USDA Forest Service, Davis, California, United States of America

Elise F. Zipkin

Affiliations USGS Patuxent Wildlife Research Center, Laurel, Maryland, United States of America, Department of Biology, University of Maryland, College Park, Maryland, United States of America

Patricia N. Manley

Affiliation Conservation of Biodiversity Group, Pacific Southwest Research Station, USDA Forest Service, Hilo, Hawaii, United States of America

Matthew D. Schlesinger

Affiliations Conservation of Biodiversity Group, Pacific Southwest Research Station, USDA Forest Service, Davis, California, United States of America, Department of Environmental Science and Policy and Graduate Group in Ecology, University of California Davis, Davis, California, United States of America, New York Natural Heritage Program, Albany, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PNM MDS. Analyzed the data: AMW EFZ. Wrote the paper: AMW EFZ PNM.