About the Authors

Zhi Zhang

Affiliation Development and Aging Program, Neuroscience, Aging and Stem Cell Research Center, Sanford-Burnham Medical Research Institute, La Jolla, California, United States of America

Benjamin Hsieh

Affiliation School of Biology, Georgia Institute of Technology, Atlanta, Georgia, United States of America

Amy Poe

Affiliation School of Biology, Georgia Institute of Technology, Atlanta, Georgia, United States of America

Julie Anderson

Affiliation Development and Aging Program, Neuroscience, Aging and Stem Cell Research Center, Sanford-Burnham Medical Research Institute, La Jolla, California, United States of America

Karen Ocorr

Affiliation Development and Aging Program, Neuroscience, Aging and Stem Cell Research Center, Sanford-Burnham Medical Research Institute, La Jolla, California, United States of America

Greg Gibson

rolf@sbmri.org (RB); greg.gibson@biology.gatech.edu (GG)

Affiliation School of Biology, Georgia Institute of Technology, Atlanta, Georgia, United States of America

Rolf Bodmer

rolf@sbmri.org (RB); greg.gibson@biology.gatech.edu (GG)

Affiliation Development and Aging Program, Neuroscience, Aging and Stem Cell Research Center, Sanford-Burnham Medical Research Institute, La Jolla, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GG RB ZZ. Performed the experiments: ZZ JA AP KO. Analyzed the data: ZZ BH KO. Contributed reagents/materials/analysis tools: ZZ BH KO. Wrote the paper: GG RB ZZ.