About the Authors

Steven H. Pearson

shp36@drexel.edu

Affiliation Drexel University, Department of Biodiversity, Earth and Environmental Science, Philadelphia, Pennsylvania, United States of America

Harold W. Avery

Affiliation Drexel University, Department of Biology, Philadelphia, Pennsylvania, United States of America

Susan S. Kilham

Affiliation Drexel University, Department of Biodiversity, Earth and Environmental Science, Philadelphia, Pennsylvania, United States of America

David J. Velinsky

Affiliation Drexel University, Department of Biodiversity, Earth and Environmental Science, Philadelphia, Pennsylvania, United States of America

James R. Spotila

Affiliation Drexel University, Department of Biodiversity, Earth and Environmental Science, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Drafted part of manuscript: DJV. Provided edits and assistance: SSK DJV JRS HWA. Conceived and designed the experiments: SHP SSK DJV JRS HWA. Performed the experiments: SHP. Analyzed the data: SHP SSK DJV JRS HWA. Contributed reagents/materials/analysis tools: SHP DJV JRS HWA. Wrote the paper: SHP.