About the Authors

Timothy J. Powell

timothy.powell@imm.ox.ac.uk (TJP); tao.dong@imm.ox.ac.uk (TD)

Affiliation MRC Human Immunology Unit, Weatherall Institute of Molecular Medicine, University of Oxford, John Radcliffe Hospital, Oxford, United Kingdom

Yanchun Peng

Affiliation MRC Human Immunology Unit, Weatherall Institute of Molecular Medicine, University of Oxford, John Radcliffe Hospital, Oxford, United Kingdom

Tamara K. Berthoud

Affiliation Jenner Institute, University of Oxford, Oxford, United Kingdom

Marie-Eve Blais

Affiliation MRC Human Immunology Unit, Weatherall Institute of Molecular Medicine, University of Oxford, John Radcliffe Hospital, Oxford, United Kingdom

Patrick J. Lillie

Affiliation Jenner Institute, University of Oxford, Oxford, United Kingdom

Adrian V. S. Hill

Affiliation Jenner Institute, University of Oxford, Oxford, United Kingdom

Sarah L. Rowland-Jones

Affiliation Jenner Institute, University of Oxford, Oxford, United Kingdom

Andrew J. McMichael

Affiliation Jenner Institute, University of Oxford, Oxford, United Kingdom

Sarah C. Gilbert

Affiliation Jenner Institute, University of Oxford, Oxford, United Kingdom

Tao Dong

timothy.powell@imm.ox.ac.uk (TJP); tao.dong@imm.ox.ac.uk (TD)

Affiliation MRC Human Immunology Unit, Weatherall Institute of Molecular Medicine, University of Oxford, John Radcliffe Hospital, Oxford, United Kingdom

Competing Interests

The authors' have read the journal's policy and have the following conflicts: SCG and AVSH are named as inventors on patents relating to methods of vaccination, including influenza vaccines: Methods and reagents for vaccination which generate a CD8+ T cell response. 1997 06 07. United States Patent Application 20110159034. Compositions and methods 2007 10 05. United States Patent Application 20100285050. All other authors have declared that no competing interests exist. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TJP AVSH SLRJ AJM SCG TD. Performed the experiments: TJP YP TKB MEB PJL TD. Analyzed the data: TJP TKB MEB PJL TD. Wrote the paper: TJP TKB SCG TD.