About the Authors

Tracy A. Nichols

tracy.a.nichols@aphis.usda.gov

Affiliation National Wildlife Research Center, US Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, Fort Collins, Colorado, United States of America

Terry R. Spraker

Affiliations Colorado State University Diagnostic Laboratory, Fort Collins, Colorado, United States of America, Prion Research Center and the Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University Prion Research Center, Fort Collins, Colorado, United States of America

Tara D. Rigg

Affiliation National Wildlife Research Center, US Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, Fort Collins, Colorado, United States of America

Crystal Meyerett-Reid

Affiliation Prion Research Center and the Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University Prion Research Center, Fort Collins, Colorado, United States of America

Clare Hoover

Affiliation Prion Research Center and the Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University Prion Research Center, Fort Collins, Colorado, United States of America

Brady Michel

Affiliation Prion Research Center and the Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University Prion Research Center, Fort Collins, Colorado, United States of America

Jifeng Bian

Affiliation Prion Research Center and the Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University Prion Research Center, Fort Collins, Colorado, United States of America

Edward Hoover

Affiliation Prion Research Center and the Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University Prion Research Center, Fort Collins, Colorado, United States of America

Thomas Gidlewski

Affiliation National Wildlife Research Center, US Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, Fort Collins, Colorado, United States of America

Aru Balachandran

Affiliation National and OIE Reference Laboratory for scrapie and chronic wasting disease, Canadian Food Inspection Agency, Ottawa, Ontario, Canada

Katherine O'Rourke

Current address: Department of Veterinary Microbiology and Pathology, College of Veterinary Medicine, Washington State University, Pullman, Washington, United States of America

Affiliation U. S. Department of Agriculture, Agricultural Research Service, Pullman, Washington, United States of America

Glenn C. Telling

Affiliation Prion Research Center and the Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University Prion Research Center, Fort Collins, Colorado, United States of America

Richard Bowen

Affiliation Department of Biomedical Sciences, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Mark D. Zabel

Contributed equally to this work with: Mark D. Zabel, Kurt C. VerCauteren

Affiliation Prion Research Center and the Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University Prion Research Center, Fort Collins, Colorado, United States of America

Kurt C. VerCauteren

Contributed equally to this work with: Mark D. Zabel, Kurt C. VerCauteren

Affiliation National Wildlife Research Center, US Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TAN TRS TG TR KV. Performed the experiments: TAN TRS TR KO JB MZ CMR CH BM RB. Analyzed the data: TAN JB KO GT MZ. Contributed reagents/materials/analysis tools: TAN TRS AB KO GT RB MZ KV EH. Wrote the paper: TAN TRS TR KO JB MZ KV.