About the Authors

Marc J. M. Rutten

Affiliation Animal Breeding and Genomics Centre, Wageningen University, Wageningen, The Netherlands

Aniek C. Bouwman

Affiliation Animal Breeding and Genomics Centre, Wageningen University, Wageningen, The Netherlands

R. Corinne Sprong

Affiliation NIZO Food Research B.V., Ede, The Netherlands

Johan A. M. van Arendonk

Affiliation Animal Breeding and Genomics Centre, Wageningen University, Wageningen, The Netherlands

Marleen H. P. W. Visker

marleen.visker@wur.nl

Affiliation Animal Breeding and Genomics Centre, Wageningen University, Wageningen, The Netherlands

Competing Interests

R. Corinne Sprong was employed by NIZO Food Research B.V. at the time of this study. This study was partly funded by the Cooperative Cattle Improvement Organization CRV. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RCS JAMA. Performed the experiments: RCS. Analyzed the data: MJMR ACB MHPWV. Wrote the paper: MJMR ACB RCS JAMA MHPWV.