About the Authors

Ying-Jiun J. Chen

nyc600@gmail.com (YJC); john.rubenstein@ucsf.edu (JLRR)

Current address: Genentech Inc, South San Francisco, California, United States of America

Affiliation Department of Psychiatry and the Nina Ireland Laboratory of Developmental Neurobiology, University of California San Francisco, San Francisco, California, United States of America

Daniel Vogt

Affiliation Department of Psychiatry and the Nina Ireland Laboratory of Developmental Neurobiology, University of California San Francisco, San Francisco, California, United States of America

Yanling Wang

Current address: Allen Institute for Brain Science, Seattle, Washington, United States of America

Affiliation Department of Psychiatry and the Nina Ireland Laboratory of Developmental Neurobiology, University of California San Francisco, San Francisco, California, United States of America

Axel Visel

Affiliation Genomics Division, Lawrence Berkeley National Laboratory, Berkeley, California, and Department of Energy Joint Genome Institute, Walnut Creek, California, United States of America

Shanni N. Silberberg

Affiliation Department of Psychiatry and the Nina Ireland Laboratory of Developmental Neurobiology, University of California San Francisco, San Francisco, California, United States of America

Cory R. Nicholas

Affiliation Eli and Edythe Broad Center of Regeneration Medicine and Stem Cell Research, University of California San Francisco, San Francisco, California, United States of America

Teruko Danjo

Affiliation Neurogenesis and Organogenesis Group, RIKEN Center for Developmental Biology, Kobe, Japan

Joshua L. Pollack

Affiliation Lung Biology Center, Department of Medicine, University of California San Francisco, San Francisco, California, United States of America

Len A. Pennacchio

Affiliation Genomics Division, Lawrence Berkeley National Laboratory, Berkeley, California, and Department of Energy Joint Genome Institute, Walnut Creek, California, United States of America

Stewart Anderson

Affiliation 3Children's Hospital of Philadelphia, University of Pennsylvania, School of Medicine, Philadelphia, Pennsylvania, United States of America

Yoshiki Sasai

Affiliation Neurogenesis and Organogenesis Group, RIKEN Center for Developmental Biology, Kobe, Japan

Scott C. Baraban

Affiliation Department of Neurological Surgery, University of California San Francisco, San Francisco, California, United States of America

Arnold R. Kriegstein

Affiliation Eli and Edythe Broad Center of Regeneration Medicine and Stem Cell Research, University of California San Francisco, San Francisco, California, United States of America

Arturo Alvarez-Buylla

Affiliation Department of Neurosurgery and Eli and Edythe Broad Center for Regeneration Medicine and Stem Cell Research, University of California San Francisco, San Francisco, California, United States of America

John L. R. Rubenstein

nyc600@gmail.com (YJC); john.rubenstein@ucsf.edu (JLRR)

Affiliation Department of Psychiatry and the Nina Ireland Laboratory of Developmental Neurobiology, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

JLRR, AAB, ARK, SCB, and CRN have affiliations with Neurona, Inc. However, this does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: YJC DV YS SCB ARK AAB JLRR CRN. Performed the experiments: YJC DV YW AV SNS. Analyzed the data: YJC JLRR JLP. Contributed reagents/materials/analysis tools: TD LAP SA YS. Wrote the paper: YJC DV AAB JLRR.