About the Authors

Tanya Golubchik

Contributed equally to this work with: Tanya Golubchik, Elizabeth M. Batty

Affiliation Department of Statistics, University of Oxford, Oxford, Oxfordshire, United Kingdom

Elizabeth M. Batty

Contributed equally to this work with: Tanya Golubchik, Elizabeth M. Batty

Affiliation Department of Statistics, University of Oxford, Oxford, Oxfordshire, United Kingdom

Ruth R. Miller

Affiliations Experimental Medicine Division, Nuffield Department of Clinical Medicine, University of Oxford, Oxford, Oxfordshire, United Kingdom, National Institute for Health Research Oxford Biomedical Research Centre, Oxford, Oxfordshire, United Kingdom

Helen Farr

Affiliations Experimental Medicine Division, Nuffield Department of Clinical Medicine, University of Oxford, Oxford, Oxfordshire, United Kingdom, National Institute for Health Research Oxford Biomedical Research Centre, Oxford, Oxfordshire, United Kingdom

Bernadette C. Young

Affiliation Experimental Medicine Division, Nuffield Department of Clinical Medicine, University of Oxford, Oxford, Oxfordshire, United Kingdom

Hanna Larner-Svensson

Affiliations National Institute for Health Research Oxford Biomedical Research Centre, Oxford, Oxfordshire, United Kingdom, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, Oxfordshire, United Kingdom

Rowena Fung

Affiliations Experimental Medicine Division, Nuffield Department of Clinical Medicine, University of Oxford, Oxford, Oxfordshire, United Kingdom, National Institute for Health Research Oxford Biomedical Research Centre, Oxford, Oxfordshire, United Kingdom

Heather Godwin

Affiliation Oxford University Hospitals National Health Service Trust, Oxford, Oxfordshire, United Kingdom

Kyle Knox

Affiliation Department of Primary Care Health Sciences, University of Oxford, Oxford, Oxfordshire, United Kingdom

Antonina Votintseva

Affiliations Experimental Medicine Division, Nuffield Department of Clinical Medicine, University of Oxford, Oxford, Oxfordshire, United Kingdom, National Institute for Health Research Oxford Biomedical Research Centre, Oxford, Oxfordshire, United Kingdom

Richard G. Everitt

Affiliation Experimental Medicine Division, Nuffield Department of Clinical Medicine, University of Oxford, Oxford, Oxfordshire, United Kingdom

Teresa Street

Affiliation Department of Statistics, University of Oxford, Oxford, Oxfordshire, United Kingdom

Madeleine Cule

Affiliation Department of Statistics, University of Oxford, Oxford, Oxfordshire, United Kingdom

Camilla L. C. Ip

Affiliation Department of Statistics, University of Oxford, Oxford, Oxfordshire, United Kingdom

Xavier Didelot

Affiliation Department of Statistics, University of Oxford, Oxford, Oxfordshire, United Kingdom

Timothy E. A. Peto

Affiliations Experimental Medicine Division, Nuffield Department of Clinical Medicine, University of Oxford, Oxford, Oxfordshire, United Kingdom, National Institute for Health Research Oxford Biomedical Research Centre, Oxford, Oxfordshire, United Kingdom

Rosalind M. Harding

Affiliation Department of Zoology, University of Oxford, Oxford, Oxfordshire, United Kingdom

Daniel J. Wilson

daniel.wilson@ndm.ox.ac.uk (DJW); bowden@stats.ox.ac.uk (RB)

These authors also contributed equally to this work.

Affiliations Experimental Medicine Division, Nuffield Department of Clinical Medicine, University of Oxford, Oxford, Oxfordshire, United Kingdom, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, Oxfordshire, United Kingdom

Derrick W. Crook

These authors also contributed equally to this work.

Affiliations Experimental Medicine Division, Nuffield Department of Clinical Medicine, University of Oxford, Oxford, Oxfordshire, United Kingdom, National Institute for Health Research Oxford Biomedical Research Centre, Oxford, Oxfordshire, United Kingdom

Rory Bowden

daniel.wilson@ndm.ox.ac.uk (DJW); bowden@stats.ox.ac.uk (RB)

These authors also contributed equally to this work.

Affiliations Department of Statistics, University of Oxford, Oxford, Oxfordshire, United Kingdom, National Institute for Health Research Oxford Biomedical Research Centre, Oxford, Oxfordshire, United Kingdom, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, Oxfordshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TG HF DWC RB. Performed the experiments: RRM RF HG KK TEAP HF HL-S AV TS. Analyzed the data: TG EMB RRM MC CLCI XD BCY RGE RMH DJW RB. Wrote the paper: TG EMB DJW RB XD TEAP DWC.