About the Authors

Kihyuck Kwak

Affiliation Department of Pathology, Johns Hopkins University, Baltimore, Maryland, United States of America

Rosie Jiang

Affiliation Department of Pathology, Johns Hopkins University, Baltimore, Maryland, United States of America

Subhashini Jagu

Affiliation Department of Pathology, Johns Hopkins University, Baltimore, Maryland, United States of America

Joshua W. Wang

Affiliation Department of Pathology, Johns Hopkins University, Baltimore, Maryland, United States of America

Chenguang Wang

Affiliation Department of Biostatistics, Johns Hopkins University, Baltimore, Maryland, United States of America

Neil D. Christensen

Affiliation Departments of Pathology, Microbiology and Immunology, Penn State University, Hershey, Pennsylvania, United States of America

Richard B. S. Roden

roden@jhmi.edu

Affiliation Department of Pathology, Johns Hopkins University, Baltimore, Maryland, United States of America

Competing Interests

The authors have read the journal's policy and have the following conflicts. Subhashini Jagu and Richard Roden are co-inventors on L2 patents (20090047301 Papillomavirus L2 N-Terminal Peptides for the Induction of Broadly Cross-Neutralizing Antibodies and 20100297144 MULTITYPE HPV PEPTIDE COMPOSITIONS AND METHODS FOR TREATMENT OR PREVENTION OF HUMAN PAPILLOMAVIRUS INFECTION) licensed to Shantha Biotechnics Ltd., GlaxoSmithKline, PaxVax, Inc. and Acambis, Inc and have received research funding from Shantha Biotechnics Ltd., Sanofi Pasteur and GlaxoSmithKline and Richard Roden is a founder of Papivax LLC. The terms of these arrangements are managed by Johns Hopkins University in accordance with its conflict of interest policies. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KK, NDC, RBSR. Performed the experiments: KK, RJ, SJ, JWW. Analyzed the data: CW, KK, RJ, SJ, JWW, RBSR. Contributed reagents/materials/analysis tools: KK, CW, NDC, RBSR. Wrote the paper: KK, CW, NDC, RBSR.