About the Authors

Susan M. Gribble

Contributed equally to this work with: Susan M. Gribble, Frances K. Wiseman

smg@sanger.ac.uk

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Frances K. Wiseman

Contributed equally to this work with: Susan M. Gribble, Frances K. Wiseman

Affiliation Department of Neurodegenerative Disease, UCL Institute of Neurology, Queen Square, London, United Kingdom

Stephen Clayton

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Elena Prigmore

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Elizabeth Langley

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Fengtang Yang

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Sean Maguire

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Beiyuan Fu

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Diana Rajan

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Olivia Sheppard

Affiliation Department of Neurodegenerative Disease, UCL Institute of Neurology, Queen Square, London, United Kingdom

Carol Scott

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Heidi Hauser

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Philip J. Stephens

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Lucy A. Stebbings

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Bee Ling Ng

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Tomas Fitzgerald

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Michael A. Quail

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Ruby Banerjee

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Kai Rothkamm

Affiliation Health Protection Agency Centre for Radiation, Chemical & Environmental Hazards Chilton, Didcot, Oxon, United Kingdom

Victor L. J. Tybulewicz

Affiliation MRC National Institute for Medical Research, The Ridgeway, Mill Hill, London, United Kingdom

Elizabeth M. C. Fisher

Affiliation Department of Neurodegenerative Disease, UCL Institute of Neurology, Queen Square, London, United Kingdom

Nigel P. Carter

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FW SMG FY EF NC VT. Performed the experiments: SMG FW EP EL SM BF DR OS BLN RB MQ. Analyzed the data: SMG SC EP EL FY CS TF. Contributed reagents/materials/analysis tools: FW SC FY CS HH PS LS BLN TF MQ. Wrote the paper: SMG FW EF KR NC VT.