About the Authors

Simon A. Black

S.Black@kent.ac.uk

Affiliation Durrell Institute of Conservation and Ecology, School of Anthropology and Conservation, University of Kent, Canterbury, Kent, United Kingdom

Amina Fellous

Affiliation Agence Nationale pour la Conservation de la Nature, Algiers, Algeria

Nobuyuki Yamaguchi

Affiliation Department of Biological and Environmental Sciences, University of Qatar, Doha, Qatar

David L. Roberts

Affiliation Durrell Institute of Conservation and Ecology, School of Anthropology and Conservation, University of Kent, Canterbury, Kent, United Kingdom

Competing Interests

Co-author David L Roberts is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SAB AF NY DLR. Performed the experiments: SAB AF DLR. Analyzed the data: SAB DLR. Contributed reagents/materials/analysis tools: SAB AF DLR. Wrote the paper: SAB AF NY DLR.