About the Authors

Mayuko Koreishi

Affiliation The Graduate School of Natural Science and Technology, Okayama University, Okayama, Japan

Thomas J. Gniadek

Affiliation Department of Pathology, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Sidney Yu

Affiliation School of Biomedical Sciences and Epithelial Cell Biology Research Center, The Chinese University of Hong Kong, Shatin, N.T., Hong Kong SAR, People’s Republic of China

Junko Masuda

Affiliation Mucosal Immunity Section, Laboratory of Host Defenses, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Yasuko Honjo

Affiliation The Research Core for Interdisciplinary Sciences (RCIS), Okayama University, Okayama, Japan

Ayano Satoh

ayano113@cc.okayama-u.ac.jp

Affiliation The Graduate School of Natural Science and Technology, Okayama University, Okayama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SY YH AS. Performed the experiments: MK JM AS. Analyzed the data: MK TG SY AS. Contributed reagents/materials/analysis tools: TG SY AS. Wrote the paper: TG SY YH AS.