About the Authors

Aileen Marshall

Contributed equally to this work with: Aileen Marshall, Margus Lukk

aileen.marshall@cruk.cam.ac.uk (AM); duncan.odom@cruk.cam.ac.uk (DTO)

Affiliations Cancer Research UK Cambridge Research Institute, Li Ka Shing Centre, Cambridge, United Kingdom, Department of Histopathology, Addenbrooke’s Hospital, Cambridge, United Kingdom

Margus Lukk

Contributed equally to this work with: Aileen Marshall, Margus Lukk

Affiliation Cancer Research UK Cambridge Research Institute, Li Ka Shing Centre, Cambridge, United Kingdom

Claudia Kutter

Affiliation Cancer Research UK Cambridge Research Institute, Li Ka Shing Centre, Cambridge, United Kingdom

Susan Davies

Affiliation Department of Histopathology, Addenbrooke’s Hospital, Cambridge, United Kingdom

Graeme Alexander

Affiliation Cambridge Hepatobiliary Unit, Addenbrooke’s Hospital, Cambridge, United Kingdom

Duncan T. Odom

aileen.marshall@cruk.cam.ac.uk (AM); duncan.odom@cruk.cam.ac.uk (DTO)

Affiliations Cancer Research UK Cambridge Research Institute, Li Ka Shing Centre, Cambridge, United Kingdom, Department of Oncology, Addenbrooke's Biomedical Campus, Hutchison-MRC Research Centre, Cambridge, United Kingdom

Competing Interests

The authors have declared that no competing interest exists.

Author Contributions

Conceived and designed the experiments: AM CK ML DTO GA. Performed the experiments: AM CK. Analyzed the data: AM ML SED. Contributed reagents/materials/analysis tools: DTO. Wrote the paper: AM ML CK SED GA DTO.