About the Authors

Xi Li

Contributed equally to this work with: Xi Li, Ti-Cao Zhang

Affiliation Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, School of Life Sciences, Fudan University, Shanghai, China

Ti-Cao Zhang

Contributed equally to this work with: Xi Li, Ti-Cao Zhang

Affiliation Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, School of Life Sciences, Fudan University, Shanghai, China

Qin Qiao

Affiliation Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, School of Life Sciences, Fudan University, Shanghai, China

Zhumei Ren

Affiliation College of Life Science and Technology, Shanxi University, Taiyuan, China

Jiayuan Zhao

Affiliation Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, School of Life Sciences, Fudan University, Shanghai, China

Takahiro Yonezawa

Affiliation Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, School of Life Sciences, Fudan University, Shanghai, China

Masami Hasegawa

Affiliation Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, School of Life Sciences, Fudan University, Shanghai, China

M. James C Crabbe

Affiliation Faculty of Creative Arts, Technologies and Science, Institute of Biomedical, Environmental Science and Technology, University of Bedfordshire, Luton, United Kingdom

Jianqiang Li

jianqiangl@hotmail.com (JL); yangzhong@fudan.edu.cn (YZ)

Affiliation Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan, China

Yang Zhong

jianqiangl@hotmail.com (JL); yangzhong@fudan.edu.cn (YZ)

Affiliations Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, School of Life Sciences, Fudan University, Shanghai, China, Institute of Biodiversity Science and Geobiology, Tibet University, Lhasa, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JL YZ. Performed the experiments: XL QQ TCZ. Analyzed the data: TCZ QQ XL. Contributed reagents/materials/analysis tools: ZR JZ TY MH. Wrote the paper: TCZ QQ MJCC YZ.