About the Authors

Nicola S. Lewis

nsl25@cam.ac.uk

Affiliation Department of Zoology, University of Cambridge, Cambridge, United Kingdom

Zurab Javakhishvili

Affiliation Institute of Ecology, Ilia State University, Tbilisi, Georgia

Colin A. Russell

Affiliations Department of Zoology, University of Cambridge, Cambridge, United Kingdom, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Ann Machablishvili

Affiliations Ivane Javakhishvili Tbilisi State University, Tbilisi, Georgia, Influenza Section, National Centre for Disease Control, Tbilisi, Georgia

Pascal Lexmond

Affiliation Department of Virology and National Influenza Center, Erasmus Medical Center, Rotterdam, The Netherlands

Josanne H. Verhagen

Affiliation Department of Virology and National Influenza Center, Erasmus Medical Center, Rotterdam, The Netherlands

Oanh Vuong

Affiliation Department of Virology and National Influenza Center, Erasmus Medical Center, Rotterdam, The Netherlands

Tinatin Onashvili

Affiliation Laboratory of the Ministry of Agriculture, Tbilisi, Georgia

Marina Donduashvili

Affiliation Laboratory of the Ministry of Agriculture, Tbilisi, Georgia

Derek J. Smith

Affiliations Department of Zoology, University of Cambridge, Cambridge, United Kingdom, Department of Virology and National Influenza Center, Erasmus Medical Center, Rotterdam, The Netherlands, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Ron A. M. Fouchier

Affiliation Department of Virology and National Influenza Center, Erasmus Medical Center, Rotterdam, The Netherlands

Competing Interests

The authors hereby declare that co-author Prof Ron A. M. Fouchier currently serves on the Editorial Board for PLOS ONE. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: NSL ZJ DJS RAMF. Performed the experiments: NSL ZJ AM PL JHV OV TO MD. Analyzed the data: NSL ZJ CAR. Contributed reagents/materials/analysis tools: NSL ZJ CAR AM TO MD DJS RAMF. Wrote the paper: NSL ZJ PL JHV RAMF.