About the Authors

Gregory V. Stark

Affiliation Biomedical Research Center, Battelle, Columbus, Ohio, United States of America

James P. Long

Affiliation Biomedical Research Center, Battelle, Columbus, Ohio, United States of America

Diana I. Ortiz

Affiliation Biomedical Research Center, Battelle, Columbus, Ohio, United States of America

Melicia Gainey

Affiliation Biomedical Research Center, Battelle, Columbus, Ohio, United States of America

Benjamin A. Carper

Affiliation Biomedical Research Center, Battelle, Columbus, Ohio, United States of America

Jingyu Feng

Affiliation Biomedical Research Center, Battelle, Columbus, Ohio, United States of America

Stephen M. Miller

Affiliation Biomedical Research Center, Battelle, Columbus, Ohio, United States of America

John E. Bigger

Affiliation Biomedical Research Center, Battelle, Columbus, Ohio, United States of America

Eric M. Vela

velae@battelle.org

Affiliation Biomedical Research Center, Battelle, Columbus, Ohio, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EMV GVS. Performed the experiments: EMV GVS JPL MG BAC JF JEB. Analyzed the data: GVS EMV JPL SMM DIO. Contributed reagents/materials/analysis tools: GVS BAC JF. Wrote the paper: GVS EMV DIO.