About the Authors

Jing Chen

Current address: Wolfson Centre for Age-Related Diseases, King's College London, London, United Kingdom

Affiliation Wellcome Trust Sanger Institute, Hinxton, Cambridgeshire, United Kingdom

Neil Ingham

Current address: Wolfson Centre for Age-Related Diseases, King's College London, London, United Kingdom

Affiliation Wellcome Trust Sanger Institute, Hinxton, Cambridgeshire, United Kingdom

Simon Clare

Affiliation Wellcome Trust Sanger Institute, Hinxton, Cambridgeshire, United Kingdom

Claire Raisen

Affiliation Wellcome Trust Sanger Institute, Hinxton, Cambridgeshire, United Kingdom

Valerie E. Vancollie

Affiliation Wellcome Trust Sanger Institute, Hinxton, Cambridgeshire, United Kingdom

Ozama Ismail

Affiliation Wellcome Trust Sanger Institute, Hinxton, Cambridgeshire, United Kingdom

Rebecca E. McIntyre

Affiliation Wellcome Trust Sanger Institute, Hinxton, Cambridgeshire, United Kingdom

Stephen H. Tsang

Affiliation Edward S. Harkness Eye Institute, Columbia University, New York, New York, United States of America

Vinit B. Mahajan

Affiliation Omics Laboratory, Department of Ophthalmology and Visual Sciences, University of Iowa, Iowa City, Iowa, United States of America

Gordon Dougan

Affiliation Wellcome Trust Sanger Institute, Hinxton, Cambridgeshire, United Kingdom

David J. Adams

Affiliation Wellcome Trust Sanger Institute, Hinxton, Cambridgeshire, United Kingdom

Jacqueline K. White

Affiliation Wellcome Trust Sanger Institute, Hinxton, Cambridgeshire, United Kingdom

Karen P. Steel

karen.steel@kcl.ac.uk

Current address: Wolfson Centre for Age-Related Diseases, King's College London, London, United Kingdom

Affiliation Wellcome Trust Sanger Institute, Hinxton, Cambridgeshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JC KPS GD. Performed the experiments: JC NI SC CR VEV OI REM SHT. Analyzed the data: JC NI SC CR VEM OI REM VBM DJA JKW KPS. Wrote the paper: JC KPS.