About the Authors

Xiaowei Yuan

Contributed equally to this work with: Xiaowei Yuan, Shizhong Zhang

Affiliation National Key Laboratory of Crop Biology, Shandong Agricultural University, Tai-An, Shandong, China

Shizhong Zhang

Contributed equally to this work with: Xiaowei Yuan, Shizhong Zhang

Affiliations National Key Laboratory of Crop Biology, Shandong Agricultural University, Tai-An, Shandong, China, National Research Center for Apple Engineering and Technology, Shandong Agricultural University, Tai-An, Shandong, China

Shiyang Liu

Affiliation National Key Laboratory of Crop Biology, Shandong Agricultural University, Tai-An, Shandong, China

Mingli Yu

Affiliation National Key Laboratory of Crop Biology, Shandong Agricultural University, Tai-An, Shandong, China

Hongyan Su

Affiliation Ludong University, Yantai, Shandong, China

Huairui Shu

Affiliations National Key Laboratory of Crop Biology, Shandong Agricultural University, Tai-An, Shandong, China, National Research Center for Apple Engineering and Technology, Shandong Agricultural University, Tai-An, Shandong, China

Xinzheng Li

lxz@sdau.edu.cn

Affiliation National Key Laboratory of Crop Biology, Shandong Agricultural University, Tai-An, Shandong, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XY SZ. Performed the experiments: XY SZ SL MY. Analyzed the data: SZ XY. Contributed reagents/materials/analysis tools: XL H. Shu H. Su. Wrote the paper: XY SZ.