About the Authors

Eva Veronesi

Affiliation VVD Program, The Pirbright Institute, Pirbright, Surrey, United Kingdom

Mark Henstock

Affiliation VVD Program, The Pirbright Institute, Pirbright, Surrey, United Kingdom

Simon Gubbins

Affiliation VVD Program, The Pirbright Institute, Pirbright, Surrey, United Kingdom

Carrie Batten

Affiliation VVD Program, The Pirbright Institute, Pirbright, Surrey, United Kingdom

Robyn Manley

Affiliation VVD Program, The Pirbright Institute, Pirbright, Surrey, United Kingdom

James Barber

Affiliation VVD Program, The Pirbright Institute, Pirbright, Surrey, United Kingdom

Bernd Hoffmann

Affiliation Institute of Diagnostic Virology, Friedrich-Loeffler-Institut, Greifswald-Insel Riems, Germany

Martin Beer

Affiliation Institute of Diagnostic Virology, Friedrich-Loeffler-Institut, Greifswald-Insel Riems, Germany

Houssam Attoui

Affiliation VVD Program, The Pirbright Institute, Pirbright, Surrey, United Kingdom

Peter Paul Clement Mertens

Affiliation VVD Program, The Pirbright Institute, Pirbright, Surrey, United Kingdom

Simon Carpenter

simon.carpenter@pirbright.ac.uk

Affiliation VVD Program, The Pirbright Institute, Pirbright, Surrey, United Kingdom

Competing Interests

The authors can confirm that Simon Gubbins, Houssam Attoui and Martin Beer are all PLOS ONE Editorial Board members. This does not alter the authors’ adherence to all the PLOS ONE policies on data sharing and materials.

Author Contributions

Conceived and designed the experiments: EV CB HA PPCM SC. Performed the experiments: EV MH RM JB. Analyzed the data: SG. Contributed reagents/materials/analysis tools: HA MH CB BH MB. Wrote the paper: SC SG EV MB BH.