About the Authors

Takeyuki Yamada

Affiliation Department of Biological Science and Technology, Faculty of Industrial Science and Technology, Tokyo University of Science, Noda-shi, Chiba, Japan

Yumiko Urano-Tashiro

Affiliation Department of Biological Science and Technology, Faculty of Industrial Science and Technology, Tokyo University of Science, Noda-shi, Chiba, Japan

Saori Tanaka

Affiliation Department of Biological Science and Technology, Faculty of Industrial Science and Technology, Tokyo University of Science, Noda-shi, Chiba, Japan

Hirotada Akiyama

Affiliation Department of Biological Science and Technology, Faculty of Industrial Science and Technology, Tokyo University of Science, Noda-shi, Chiba, Japan

Fumio Tashiro

ftashir@rs.noda.tus.ac.jp

Affiliation Department of Biological Science and Technology, Faculty of Industrial Science and Technology, Tokyo University of Science, Noda-shi, Chiba, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TY YU-T FT. Performed the experiments: TY YU-T ST HA FT. Analyzed the data: TY ST HA FT. Contributed reagents/materials/analysis tools: TY ST HA FT. Wrote the paper: TY FT.