About the Authors

Lin Dai

Contributed equally to this work with: Lin Dai, Chao Xu, Ming Tian, Jian Sang

Affiliations School of Computer Science and Technology, Beijing Institute of Technology, Beijing, China, CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China

Chao Xu

Contributed equally to this work with: Lin Dai, Chao Xu, Ming Tian, Jian Sang

Affiliation School of Computer Science and Technology, Beijing Institute of Technology, Beijing, China

Ming Tian

Contributed equally to this work with: Lin Dai, Chao Xu, Ming Tian, Jian Sang

Affiliation School of Computer Science and Technology, Beijing Institute of Technology, Beijing, China

Jian Sang

Contributed equally to this work with: Lin Dai, Chao Xu, Ming Tian, Jian Sang

Affiliation Research Institute of Subtropical Forestry, Chinese Academy of Forestry, Fuyang, Zhejiang, China

Dong Zou

Affiliation CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China

Ang Li

Affiliation CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China

Guocheng Liu

Affiliation CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China

Fei Chen

Affiliation CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China

Jiayan Wu

Affiliation CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China

Jingfa Xiao

Affiliation CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China

Xumin Wang

Affiliation CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China

Jun Yu

Affiliation CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China

Zhang Zhang

zhangzhang@big.ac.cn

Affiliation CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China

Competing Interests

The co-author Zhang Zhang is a PLOS ONE Editorial Board member. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Collected the nomenclature: CX MT JS GL. Developed the website: CX MT JS DZ AL. Revised the manuscript: LD FC JW JX XW JY ZZ. Conceived and designed the experiments: JY ZZ. Wrote the paper: LD ZZ.