About the Authors

Huiren Cao

Contributed equally to this work with: Huiren Cao, Xiaolei Yin

Affiliation Shandong Provincial Key Laboratory of Animal Cells and Developmental Biology, Institute of Developmental Biology, School of Life Sciences, Shandong University, Jinan, Shandong, People’s Republic of China

Xiaolei Yin

Contributed equally to this work with: Huiren Cao, Xiaolei Yin

Affiliation Shandong Provincial Key Laboratory of Animal Cells and Developmental Biology, Institute of Developmental Biology, School of Life Sciences, Shandong University, Jinan, Shandong, People’s Republic of China

Yujie Cao

Affiliation Shandong Provincial Key Laboratory of Animal Cells and Developmental Biology, Institute of Developmental Biology, School of Life Sciences, Shandong University, Jinan, Shandong, People’s Republic of China

Yecheng Jin

Affiliation Shandong Provincial Key Laboratory of Animal Cells and Developmental Biology, Institute of Developmental Biology, School of Life Sciences, Shandong University, Jinan, Shandong, People’s Republic of China

Shan Wang

Affiliation Shandong Provincial Key Laboratory of Animal Cells and Developmental Biology, Institute of Developmental Biology, School of Life Sciences, Shandong University, Jinan, Shandong, People’s Republic of China

Yanhui Kong

Affiliation Shandong Provincial Key Laboratory of Animal Cells and Developmental Biology, Institute of Developmental Biology, School of Life Sciences, Shandong University, Jinan, Shandong, People’s Republic of China

Yuexing Chen

Affiliation Shandong Provincial Key Laboratory of Animal Cells and Developmental Biology, Institute of Developmental Biology, School of Life Sciences, Shandong University, Jinan, Shandong, People’s Republic of China

Jiangang Gao

Affiliation Shandong Provincial Key Laboratory of Animal Cells and Developmental Biology, Institute of Developmental Biology, School of Life Sciences, Shandong University, Jinan, Shandong, People’s Republic of China

Stefan Heller

Affiliation Departments of Otolaryngology – Head & Neck Surgery and Molecular & Cellular Physiology, Stanford University School of Medicine, Stanford, California, United States of America

Zhigang Xu

xuzg@sdu.edu.cn

Affiliation Shandong Provincial Key Laboratory of Animal Cells and Developmental Biology, Institute of Developmental Biology, School of Life Sciences, Shandong University, Jinan, Shandong, People’s Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SH ZX. Performed the experiments: HC XY YC YJ SW YK YC ZX. Analyzed the data: JG SH ZX. Contributed reagents/materials/analysis tools: JG. Wrote the paper: SH ZX.