About the Authors

Gregory J. Tanner

Contributed equally to this work with: Gregory J. Tanner, Michelle L. Colgrave

Greg.Tanner@csiro.au

Affiliations Commonwealth Scientific and Industrial Research Organisation Plant Industry, Canberra, Australian Capital Territory, Australia, Commonwealth Scientific and Industrial Research Organisation Food Futures Flagship, Riverside Corporate Park, North Ryde, New South Wales, Australia

Michelle L. Colgrave

Contributed equally to this work with: Gregory J. Tanner, Michelle L. Colgrave

Affiliations Commonwealth Scientific and Industrial Research Organisation Food Futures Flagship, Riverside Corporate Park, North Ryde, New South Wales, Australia, Commonwealth Scientific and Industrial Research Organisation Animal, Food and Health Sciences, St Lucia, Queensland, Australia

Malcolm J. Blundell

Affiliations Commonwealth Scientific and Industrial Research Organisation Plant Industry, Canberra, Australian Capital Territory, Australia, Commonwealth Scientific and Industrial Research Organisation Food Futures Flagship, Riverside Corporate Park, North Ryde, New South Wales, Australia

Hareshwar P. Goswami

Affiliations Commonwealth Scientific and Industrial Research Organisation Food Futures Flagship, Riverside Corporate Park, North Ryde, New South Wales, Australia, Commonwealth Scientific and Industrial Research Organisation Animal, Food and Health Sciences, St Lucia, Queensland, Australia

Crispin A. Howitt

Affiliations Commonwealth Scientific and Industrial Research Organisation Plant Industry, Canberra, Australian Capital Territory, Australia, Commonwealth Scientific and Industrial Research Organisation Food Futures Flagship, Riverside Corporate Park, North Ryde, New South Wales, Australia

Competing Interests

We have the following interests: Tanner and Howitt are inventors on a provisional patent application “Barley with low levels of hordein” U.S. Provisional application No. 505676, filed Aug 13, 2007; International Patent Application No. PCT/AU2008/001172 (WO2009021285-A1). The authors derive no financial benefit from this patent and all rights have been assigned to CSIRO, Walter & Eliza Hall Institute of Medical Research, Melbourne Health and Grains Research and Development Commission (Australia). As far as we are aware as at the time of submission of this paper, there are no further patents, commercial products in development, or marketed products related to the subject matter of this paper to declare. This does not alter our adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Designed the ELISA experiments: GT MJB. Designed the MS experements: HG MLC. Collected ELISA data: GT MJB. Carried out the MS experiments: HG MLC. Analysed the ELISA protein data: GT MJB. Analysed the MS data: HG MLC. Contributed analysis tools for the ELISA experiments: GT MJB. Contributed analysis tools for the MS data: HG MLC. Wrote the paper: GT MLB MLC HG CAH.