About the Authors

Lucas J. Beversdorf

lbeversdorf@wisc.edu

Affiliation Department of Civil and Environmental Engineering, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Todd R. Miller

Affiliation Zilber School of Public Health, University of Wisconsin-Milwaukee, Milwaukee, Wisconsin, United States of America

Katherine D. McMahon

Affiliations Department of Civil and Environmental Engineering, University of Wisconsin-Madison, Madison, Wisconsin, United States of America, Department of Bacteriology, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LJB. Performed the experiments: LJB. Analyzed the data: LJB TRM. Contributed reagents/materials/analysis tools: LJB TRM KDM. Wrote the paper: LJB TRM KDM.