About the Authors

Gil Bohrer

bohrer.17@osu.edu

Affiliation Department of Civil, Environmental and Geodetic Engineering, The Ohio State University, Columbus, Ohio, United States of America

Kunpeng Zhu

Affiliation Department of Civil, Environmental and Geodetic Engineering, The Ohio State University, Columbus, Ohio, United States of America

Robert L. Jones

Affiliation Department of Civil, Environmental and Geodetic Engineering, The Ohio State University, Columbus, Ohio, United States of America

Peter S. Curtis

Affiliation Department of, Evolution, Ecology, and Organismal Biology, The Ohio State University, Columbus, Ohio, United States of America

Competing Interests

Gil Bohrer is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials. Gil Bohrer did not participate in any way in the editorial process of this manuscript.

Author Contributions

Conceived and designed the experiments: GB PSC. Performed the experiments: KZ RLJ. Analyzed the data: KZ GB. Contributed reagents/materials/analysis tools: GB. Wrote the paper: GB KZ PSC.