About the Authors

Gretel Torres de la Riva

Affiliation Department of Population Health and Reproduction, School of Veterinary Medicine, University of California-Davis, Davis, California, United States of America,

Benjamin L. Hart

blhart@ucdavis.edu

Affiliation Department of Anatomy, Physiology and Cell Biology, School of Veterinary Medicine, University of California-Davis, Davis, California, United States of America

Thomas B. Farver

Affiliation Department of Population Health and Reproduction, School of Veterinary Medicine, University of California-Davis, Davis, California, United States of America,

Anita M. Oberbauer

Affiliation Department of Animal Science, College of Agriculture and Environmental Sciences, University of California-Davis, Davis, California, United States of America

Locksley L. McV. Messam

Affiliation Department of Public Health Sciences, School of Medicine, University of California-Davis, Davis, California, United States of America

Neil Willits

Affiliation Statistics Laboratory, Department of Statistics, University of California-Davis, Davis, California, United States of America

Lynette A. Hart

Affiliation Department of Population Health and Reproduction, School of Veterinary Medicine, University of California-Davis, Davis, California, United States of America,

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Interpreting results and editing manuscript: AMO. Conceived and designed the experiments: BLH LAH GT. Performed the experiments: GT LAH BLH. Analyzed the data: GT TBF LLM NW. Wrote the paper: GT BLH LAH.