About the Authors

Michael P. Heaton

Contributed equally to this work with: Michael P. Heaton, Theodore S. Kalbfleisch

mike.heaton@ars.usda.gov (MPH); ted.kalbfleisch@louisville.edu (TSK)

Affiliation U.S. Meat Animal Research Center (USMARC), Clay Center, Nebraska, United States of America

Theodore S. Kalbfleisch

Contributed equally to this work with: Michael P. Heaton, Theodore S. Kalbfleisch

mike.heaton@ars.usda.gov (MPH); ted.kalbfleisch@louisville.edu (TSK)

Affiliations Department of Biochemistry and Molecular Biology, School of Medicine, University of Louisville, Louisville, Kentucky, United States of America, Intrepid Bioinformatics, Louisville, Kentucky, United States of America

Dustin T. Petrik

Affiliation GeneSeek, a Neogen company, Lincoln, Nebraska, United States of America

Barry Simpson

Affiliation GeneSeek, a Neogen company, Lincoln, Nebraska, United States of America

James W. Kijas

Affiliation Division of Animal, Food and Health Sciences, CSIRO, Brisbane, Australia

Michael L. Clawson

Affiliation U.S. Meat Animal Research Center (USMARC), Clay Center, Nebraska, United States of America

Carol G. Chitko-McKown

Affiliation U.S. Meat Animal Research Center (USMARC), Clay Center, Nebraska, United States of America

Gregory P. Harhay

Affiliation U.S. Meat Animal Research Center (USMARC), Clay Center, Nebraska, United States of America

Kreg A. Leymaster

Affiliation U.S. Meat Animal Research Center (USMARC), Clay Center, Nebraska, United States of America

the International Sheep Genomics Consortium

Membership of the International Sheep Genomics Consortium is provided in the Acknowledgments and at http://www.sheephapmap.org/participants.php.

Competing Interests

The authors have the following interests: co-authors D.T.P. and B.S. are full-time employees at GeneSeek, a Neogen company that provides agrigenomic and veterinary diagnostic services. T.S.K. is the CEO of Intrepid Bioinformatics, a company that provides web-based systems to privately store, analyze, curate, share, and remotely access genetic data. International Sheep Genomics Consortium member K.G. is employed by Illumina Inc. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Performed the experiments: MPH TSK DTP BS. Conceived and designed the experiments: MPH TSK DTP BS MLC CGC-M GPH KAL. Analyzed the data: MPH TSK DTP BS KAL. Wrote the paper: MPH TSK DTP JWK MLC CGC-M GPH KAL. Contributed reagents/materials/analysis tools: MPH TSK DTP BS JWK KAL ISGC.