About the Authors

Kylee L. Spencer

Contributed equally to this work with: Kylee L. Spencer, Jennifer Malinowski, Steven Buyske, Dana C. Crawford

Affiliation Department of Biology and Environmental Science, Heidelberg University, Tiffin, Ohio, United States of America

Jennifer Malinowski

Contributed equally to this work with: Kylee L. Spencer, Jennifer Malinowski, Steven Buyske, Dana C. Crawford

Affiliation Center for Human Genetics Research, Department of Molecular Physiology and Biophysics, Vanderbilt University, Nashville, Tennessee, United States of America

Cara L. Carty

Affiliation Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Nora Franceschini

Affiliation Department of Epidemiology, University of North Carolina, Chapel Hill, North Carolina, United States of America

Lindsay Fernández-Rhodes

Affiliation Department of Epidemiology, University of North Carolina, Chapel Hill, North Carolina, United States of America

Alicia Young

Affiliation Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Iona Cheng

Affiliation University of Hawaii Cancer Center, University of Hawaii, Honolulu, Hawaii, United States of America

Marylyn D. Ritchie

Affiliations Center for Human Genetics Research, Department of Molecular Physiology and Biophysics, Vanderbilt University, Nashville, Tennessee, United States of America, Biochemistry and Molecular Biology, Penn State University, University Park, Maryland, United States of America

Christopher A. Haiman

Affiliation Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

Lynne Wilkens

Affiliation University of Hawaii Cancer Center, University of Hawaii, Honolulu, Hawaii, United States of America

ChunyuanWu

Affiliation Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Tara C. Matise

Affiliation Department of Genetics, Rutgers University, Piscataway, New Jersey, United States of America

Christopher S. Carlson

Affiliation Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Kathleen Brennan

Affiliation Department of Obstetrics and Gynecology, University of California Los Angeles, Los Angeles, California, United States of America

Amy Park

Affiliation Department of Obstetrics and Gynecology, School of Medicine, Georgetown University, Washington, DC, United States of America

Aleksandar Rajkovic

Affiliation Magee-Womens Research Institute, Department of Obstetrics, Gynecology and Reproductive Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Lucia A. Hindorff

Affiliation Office of Population Genomics, National Human Genome Research Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Steven Buyske

Contributed equally to this work with: Kylee L. Spencer, Jennifer Malinowski, Steven Buyske, Dana C. Crawford

Affiliations Department of Genetics, Rutgers University, Piscataway, New Jersey, United States of America, Department of Statistics, Rutgers University, Piscataway, New Jersey, United States of America

Dana C. Crawford

Contributed equally to this work with: Kylee L. Spencer, Jennifer Malinowski, Steven Buyske, Dana C. Crawford

dana.c.crawford@vanderbilt.edu

Affiliation Center for Human Genetics Research, Department of Molecular Physiology and Biophysics, Vanderbilt University, Nashville, Tennessee, United States of America

Competing Interests

Co-author Dana C. Crawford is a PLOS ONE Editorial Board member. This membership does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KLS CLC NF LFR AY IC MDR CAH LW CW TCM CSC KB AP AR LAH SB DCC. Performed the experiments: KLS CLC NF LFR AY IC MDR CAH LW CW TCM CSC KB AP AR LAH SB DCC. Analyzed the data: KLS JM CLC NF LFR AY IC MDR CAH LW CW TCM CSC KB AP AR LAH SB DCC. Contributed reagents/materials/analysis tools: KLS JM CLC NF LFR AY IC MDR CAH LW CW TCM CSC KB AP AR SB DCC. Wrote the paper: KLS JM CLC NF LFR LAH SB DCC.