About the Authors

Rosemarie Seymour

rosemarie.seymour@gmail.com

Current address: American Veterinary Medical Association, Schaumburg, Illinois, United States of America

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Bobbi-Jo Shirley

Current address: Applied Spectral Imaging, Carlsbad, California, United States of America

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Harm HogenEsch

Affiliation Department of Comparative Pathobiology, Purdue University, West Lafayette, Indiana, United States of America

Leonard D. Shultz

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

John P. Sundberg

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Performed the experiments: BJS RES JPS LDS. Conceived and designed the experiments: RES BJS JPS. Intellectual contributions, critical revision of manuscript: LDS JPS HH RES. Wrote the paper: RES HH JPS LDS. Analyzed the data: RES BJS HH JPS. Contributed reagents/materials/analysis tools: RES JPS LDS.