About the Authors

Yuriko Matsuzaki

Affiliation Division of Gene Regulation, Institute for Advanced Medical Research, School of Medicine, Keio University, Tokyo, Japan

Haru Hosokai

Affiliation Division of Material and Biological Sciences, Japan Women's University, Tokyo, Japan

Yukiyo Mizuguchi

Affiliation Division of Gene Regulation, Institute for Advanced Medical Research, School of Medicine, Keio University, Tokyo, Japan

Shoji Fukamachi

Affiliation Laboratory of Evolutionary Genetics, Department of Chemical and Biological Sciences, Japan Women's University, Tokyo, Japan

Atsushi Shimizu

Affiliation Department of Molecular Biology, School of Medicine, Keio University, Tokyo, Japan

Hideyuki Saya

hsaya@a5.keio.jp

Affiliations Division of Gene Regulation, Institute for Advanced Medical Research, School of Medicine, Keio University, Tokyo, Japan, Japan Science and Technology Agency, Core Research for Evolutional Science and Technology, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: Y. Matsuzaki HS. Performed the experiments: Y. Matsuzaki HH Y. Mizuguchi AS. Analyzed the data: Y. Matsuzaki HH HS SF HS. Contributed reagents/materials/analysis tools: Y. Matsuzaki HH Y. Mizuguchi AS SF. Wrote the paper: Y. Matsuzaki HS.