About the Authors

Zhang Liping

Affiliations Jiangsu Key Lab of Marine Biology, College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing, China, Key Laboratory of Coastal Biology & Bioresources Utilization, Yantai Institute of Costal Zone Research(YIC), Chinese Academy of Sciences (CAS), Yantai, China

Shao Hongbo

shaohongbochu@126.com (SHB); longxiaohua@njau.edu.cn (LXH); sea@njau.edu.cn (LZP)

Affiliations Jiangsu Key Lab of Marine Biology, College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing, China, Key Laboratory of Coastal Biology & Bioresources Utilization, Yantai Institute of Costal Zone Research(YIC), Chinese Academy of Sciences (CAS), Yantai, China, Institute of Life Sciences, Qingdao University of Science and Technology, Qingdao, China

Long Xiaohua

shaohongbochu@126.com (SHB); longxiaohua@njau.edu.cn (LXH); sea@njau.edu.cn (LZP)

Affiliation Jiangsu Key Lab of Marine Biology, College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing, China

Liu Zhaopu

shaohongbochu@126.com (SHB); longxiaohua@njau.edu.cn (LXH); sea@njau.edu.cn (LZP)

Affiliation Jiangsu Key Lab of Marine Biology, College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LZP. Performed the experiments: ZPL. Analyzed the data: SHB LZP LXH. Contributed reagents/materials/analysis tools: LZP LXH. Wrote the paper: ZLP SHB LXH.