About the Authors

Mary H. Young

Affiliation Integrated Department of Immunology, National Jewish Health and University of Colorado School of Medicine, Denver, Colorado, United States of America

Lance U’Ren

Affiliation Integrated Department of Immunology, National Jewish Health and University of Colorado School of Medicine, Denver, Colorado, United States of America

Shouxiong Huang

Current address: Division of Rheumatology, Immunology and Allergy, Brigham and Women’s Hospital, Harvard Medical School, Boston, Massachusetts, United States of America

Affiliation Department of Pathology and Immunology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Thierry Mallevaey

Current address: University of Toronto, Toronto, Ontario, Canada

Affiliation Integrated Department of Immunology, National Jewish Health and University of Colorado School of Medicine, Denver, Colorado, United States of America

James Scott-Browne

Affiliation Integrated Department of Immunology, National Jewish Health and University of Colorado School of Medicine, Denver, Colorado, United States of America

Frances Crawford

Affiliation Integrated Department of Immunology, National Jewish Health and University of Colorado School of Medicine, Denver, Colorado, United States of America

Olivier Lantz

Affiliation Inserm Unit 732, Institut Curie, Paris, France

Ted H. Hansen

Affiliation Department of Pathology and Immunology, Washington University School of Medicine, St. Louis, Missouri, United States of America

John Kappler

Affiliations Integrated Department of Immunology, National Jewish Health and University of Colorado School of Medicine, Denver, Colorado, United States of America, Howard Hughes Medical Institute, University of Colorado School of Medicine, Aurora, Colorado, United States of America, Department of Medicine, University of Colorado School of Medicine, Aurora, Colorado, United States of America, Department of Pharmacology, University of Colorado School of Medicine, Aurora, Colorado, United States of America, Program in Biomolecular Structure, University of Colorado School of Medicine, Aurora, Colorado, United States of America

Philippa Marrack

Contributed equally to this work with: Philippa Marrack, Laurent Gapin

gapinl@njhealth.org (LG); marrackp@njhealth.org (PM)

Affiliations Integrated Department of Immunology, National Jewish Health and University of Colorado School of Medicine, Denver, Colorado, United States of America, Howard Hughes Medical Institute, University of Colorado School of Medicine, Aurora, Colorado, United States of America, Department of Medicine, University of Colorado School of Medicine, Aurora, Colorado, United States of America, Department of Biochemistry and Molecular Genetics, University of Colorado School of Medicine, Aurora, Colorado, United States of America

Laurent Gapin

Contributed equally to this work with: Philippa Marrack, Laurent Gapin

gapinl@njhealth.org (LG); marrackp@njhealth.org (PM)

Affiliation Integrated Department of Immunology, National Jewish Health and University of Colorado School of Medicine, Denver, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MHY LU PM LG. Performed the experiments: MHY LU. Analyzed the data: TM JS-B FC SH OL THH JK PM LG. Contributed reagents/materials/analysis tools: SH OL THH. Wrote the paper: MHY THH OL PM LG.