About the Authors

Timothy S. Frana

tfrana@iastate.edu

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Aleigh R. Beahm

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Blake M. Hanson

Affiliation University of Iowa, Center for Emerging Infectious Diseases, Department of Epidemiology, College of Public Health, Iowa City, Iowa, United States of America

Joann M. Kinyon

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Lori L. Layman

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Locke A. Karriker

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Alejandro Ramirez

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Tara C. Smith

Affiliation University of Iowa, Center for Emerging Infectious Diseases, Department of Epidemiology, College of Public Health, Iowa City, Iowa, United States of America

Competing Interests

TS is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TSF ARB JMK LAK AR TCS. Performed the experiments: ARB BMH JMK LLL LAK AR. Analyzed the data: TSF ARB BMH. Contributed reagents/materials/analysis tools: TSF TCS. Wrote the paper: TSF ARB.