About the Authors

Makoto Nasu

Affiliation Organogenesis and Neurogenesis Group, Center for Developmental Biology, RIKEN, Kobe, Japan

Nozomu Takata

Affiliation Organogenesis and Neurogenesis Group, Center for Developmental Biology, RIKEN, Kobe, Japan

Teruko Danjo

Affiliation Organogenesis and Neurogenesis Group, Center for Developmental Biology, RIKEN, Kobe, Japan

Hideya Sakaguchi

Affiliation Organogenesis and Neurogenesis Group, Center for Developmental Biology, RIKEN, Kobe, Japan

Taisuke Kadoshima

Affiliations Organogenesis and Neurogenesis Group, Center for Developmental Biology, RIKEN, Kobe, Japan, Four-Dimensional Tissue Analysis Unit, Center for Developmental Biology, RIKEN, Kobe, Japan

Sugiko Futaki

Affiliation Laboratory of Extracellular Matrix Biochemistry, Institute for Protein Research, Osaka University, Suita, Japan

Kiyotoshi Sekiguchi

Affiliation Laboratory of Extracellular Matrix Biochemistry, Institute for Protein Research, Osaka University, Suita, Japan

Mototsugu Eiraku

Affiliation Four-Dimensional Tissue Analysis Unit, Center for Developmental Biology, RIKEN, Kobe, Japan

Yoshiki Sasai

yoshikisasai@cdb.riken.jp

Affiliations Organogenesis and Neurogenesis Group, Center for Developmental Biology, RIKEN, Kobe, Japan, Four-Dimensional Tissue Analysis Unit, Center for Developmental Biology, RIKEN, Kobe, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MN YS. Performed the experiments: MN NT TD HS TK ME. Analyzed the data: MN ME. Contributed reagents/materials/analysis tools: SF KS. Wrote the paper: MN YS.