About the Authors

Asim K. Bepari

Contributed equally to this work with: Asim K. Bepari, Hiromi Sano

Affiliations Department of Morphological Neural Science, Graduate School of Medical Sciences, Kumamoto University, Kumamoto, Japan, Division of Neurobiology and Anatomy, Graduate School of Medical and Dental Sciences, Niigata University, Niigata, Japan

Hiromi Sano

Contributed equally to this work with: Asim K. Bepari, Hiromi Sano

Affiliation Division of System Neurophysiology, National Institute for Physiological Sciences, Okazaki, Japan

Nobuaki Tamamaki

Affiliation Department of Morphological Neural Science, Graduate School of Medical Sciences, Kumamoto University, Kumamoto, Japan

Atsushi Nambu

Affiliations Division of System Neurophysiology, National Institute for Physiological Sciences, Okazaki, Japan, Department of Physiological Sciences, Graduate University for Advanced Studies (Sokendai), Okazaki, Japan

Kenji F. Tanaka

kftanaka@a8.keio.jp (KT); takebaya@med.niigata-u.ac.jp (HT)

Affiliation Department of Neuropsychiatry, Keio University School of Medicine, Tokyo, Japan

Hirohide Takebayashi

kftanaka@a8.keio.jp (KT); takebaya@med.niigata-u.ac.jp (HT)

Affiliations Division of Neurobiology and Anatomy, Graduate School of Medical and Dental Sciences, Niigata University, Niigata, Japan, PRESTO, Japan Science and Technology Agency (JST), Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AKB KFT HT. Performed the experiments: AKB HS KFT. Analyzed the data: AKB KFT HT. Contributed reagents/materials/analysis tools: AKB HS NT AN KFT HT. Wrote the paper: AKB KFT HT.