About the Authors

Shuo Jiao

sjiao@fhcrc.org

Affiliation Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Li Hsu

Affiliation Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Sonja Berndt

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, Bethesda, Maryland, United States of America

Stéphane Bézieau

Affiliation Service de Génétique Médicale, CHU Nantes, Nantes, France

Hermann Brenner

Affiliation Division of Clinical Epidemiology and Aging Research, German Cancer Research Center, Heidelberg, Germany

Daniel Buchanan

Affiliation Cancer and Population Studies Group, Queensland Institute of Medical Research, Queensland, Australia

Bette J. Caan

Affiliation Division of Research, Kaiser Permanente Medical Care Program, Oakland, California, United States of America

Peter T. Campbell

Affiliation Epidemiology Research Program, American Cancer Society, Atlanta, Georgia, United States of America

Christopher S. Carlson

Affiliations Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, School of Public Health, University of Washington, Seattle, Washington, United States of America

Graham Casey

Affiliation Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

Andrew T. Chan

Affiliations Division of Gastroenterology, Massachusetts General Hospital and Harvard Medical School, Boston, Massachusetts, United States of America, Channing Division of Network Medicine, Brigham and Women’s Hospital and Harvard Medical School, Boston, Massachusetts, United States of America

Jenny Chang-Claude

Affiliation Division of Cancer Epidemiology, German Cancer Research Center, Heidelberg, Germany

Stephen Chanock

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, Bethesda, Maryland, United States of America

David V. Conti

Affiliation Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

Keith R. Curtis

Affiliation Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

David Duggan

Affiliation Translational Genomics Research Institute, Phoenix, Arizona, United States of America

Steven Gallinger

Affiliation Department of Surgery, University Health Network, Toronto General Hospital, Toronto, Canada

Stephen B. Gruber

Affiliation Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

Tabitha A. Harrison

Affiliation Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Richard B. Hayes

Affiliation Division of Epidemiology, New York University School of Medicine, New York, New York, United States of America

Brian E. Henderson

Affiliation Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

Michael Hoffmeister

Affiliation Division of Clinical Epidemiology and Aging Research, German Cancer Research Center, Heidelberg, Germany

John L. Hopper

Affiliation Melbourne School of Population Health, University of Melbourne, Melbourne, Australia

Thomas J. Hudson

Affiliations Ontario Institute for Cancer Research, Toronto, Canada, Departments of Medical Biophysics and Molecular Genetics, University of Toronto, Toronto, Canada

Carolyn M. Hutter

Affiliation Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Rebecca D. Jackson

Affiliation Division of Endocrinology, Diabetes, and Metabolism, Ohio State University, Columbus, Ohio, United States of America

Mark A. Jenkins

Affiliation Melbourne School of Population Health, University of Melbourne, Melbourne, Australia

Elizabeth D. Kantor

Affiliation Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Laurence N. Kolonel

Affiliation Epidemiology Program, University of Hawaii Cancer Center, Honolulu, Hawaii, United States of America

Sébastien Küry

Affiliation Service de Génétique Médicale, CHU Nantes, Nantes, France

Loic Le Marchand

Affiliation Epidemiology Program, University of Hawaii Cancer Center, Honolulu, Hawaii, United States of America

Mathieu Lemire

Affiliation Ontario Institute for Cancer Research, Toronto, Canada

Polly A. Newcomb

Affiliations Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, School of Public Health, University of Washington, Seattle, Washington, United States of America

John D. Potter

Affiliations Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, School of Public Health, University of Washington, Seattle, Washington, United States of America, Centre for Public Health Research, Massey University, Wellington, New Zealand

Conghui Qu

Affiliation Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Stephanie A. Rosse

Affiliation Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Robert E. Schoen

Affiliation Department of Medicine and Epidemiology, University of Pittsburgh Medical Center, Pittsburgh, Pennsylvania, United States of America

Fred R. Schumacher

Affiliation Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

Daniela Seminara

Affiliation Division of Cancer Control and Population Sciences, National Cancer Institute, Bethesda, Maryland, United States of America

Martha L. Slattery

Affiliation Department of Internal Medicine, University of Utah Health Sciences Center, Salt Lake City, Utah, United States of America

Cornelia M. Ulrich

Affiliations Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, School of Public Health, University of Washington, Seattle, Washington, United States of America, Division of Preventive Oncology, National Center for Tumor Diseases and German Cancer Research Center, Heidelberg, Germany

Brent W. Zanke

Affiliation Clinical Epidemiology Program, Ottawa Hospital Research Institute, Ottawa, Canada

Ulrike Peters

Affiliations Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, School of Public Health, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SJ LH SB SB HB DB BJC CSC GC ATC JC SC DVC DD SG SBG RBH BEH MH JLH TJH CMH RDJ MAJ LNK SK LL ML PAN JDP RES FRS DS MLS CMU BWZ UP. Performed the experiments: SJ LH SB SB HB DB BJC CSC GC ATC JC SC DVC KRC DD SG SBG TAH RBH BEH MH JLH TJH CMH RDJ MAJ EDK LNK SK LL ML PAN JDP CQ RES FRS DS MLS CMU BWZ UP. Analyzed the data: SJ LH KRC CMH CQ. Contributed reagents/materials/analysis tools: SJ LH CMH CQ UP SAR KRC PTC. Wrote the paper: SJ LH CMH UP SAR MAJ ML EDK JDP CMU.