About the Authors

Charlotte Tollenaere

Affiliation Metapopulation Research Group, Department of Biological and Environmental Sciences, University of Helsinki, Helsinki, Finland

Hanna Susi

Affiliation Metapopulation Research Group, Department of Biological and Environmental Sciences, University of Helsinki, Helsinki, Finland

Jussi Nokso-Koivisto

Affiliation Metapopulation Research Group, Department of Biological and Environmental Sciences, University of Helsinki, Helsinki, Finland

Patrik Koskinen

Affiliations Metapopulation Research Group, Department of Biological and Environmental Sciences, University of Helsinki, Helsinki, Finland, Institute of Biotechnology, University of Helsinki, Helsinki, Finland

Ayco Tack

Affiliation Metapopulation Research Group, Department of Biological and Environmental Sciences, University of Helsinki, Helsinki, Finland

Petri Auvinen

Affiliation Institute of Biotechnology, University of Helsinki, Helsinki, Finland

Lars Paulin

Affiliation Institute of Biotechnology, University of Helsinki, Helsinki, Finland

Mikko J. Frilander

Affiliation Institute of Biotechnology, University of Helsinki, Helsinki, Finland

Rainer Lehtonen

Affiliation Metapopulation Research Group, Department of Biological and Environmental Sciences, University of Helsinki, Helsinki, Finland

Anna-Liisa Laine

anna-liisa.laine@helsinki.fi

Affiliation Metapopulation Research Group, Department of Biological and Environmental Sciences, University of Helsinki, Helsinki, Finland

Competing Interests

Anna-Liisa Laine is a PLOS ONE Editorial Board member, but this does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ALL MF RL CT HS. Performed the experiments: CT HS MF PA LP AT. Analyzed the data: JNK PK CT HS. Wrote the paper: CT ALL HS.